Fireflies hibernate over winter during the larval stage, some species for several years. Some do this by burrowing underground, while others find places on or under the bark of trees. They emerge in the spring. After several weeks of feeding, they pupate for 1 to 2.5 weeks and emerge as adults. The larvae of most species are specialized predators and feed on other larvae, terrestrial snails, and slugs. Some are so specialized that they have grooved mandibles which deliver digestive fluids directly to their prey. Adult diet varies. Some are predatory, while others feed on plant pollen or nectar.
Most fireflies are quite distasteful and sometimes poisonous to vertebrate predators. This is due at least in part to a group of steroid pyrones known as lucibufagins (LBGs), which are similar to cardiotonicbufadienolides found in some poisonous toads.

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